Comment Seems like they are characterizing the sensitivity (Score 1) 63
of biological systems to mutation more than anything else. Without mutation you will just see the permutations of the existing genome try to fill the ecological space with those best suited to particular niches taking up residence there. The lizards for example the best suited fro tree climbing/feeding take up residence in the trees and breed new generations with an ever more narrow slice of the original colonizers genome.
The beneficial mutations that introduce new genes are were you get changes and would seem to be nearly impossible to predict in detail. You could hypothesize that a bacterial would eventually mutate to utilize an available food source, if not in any great detail the exact method.
The rest just seems to be usual determinism. Does it really take that much to predict that if you breed horse in pens with 3 foot ceilings and keep them in them all the time, in a few generations you are going to get short horses ?