Ever pause to consider that this may have been done intentionally as an example to demonstrate how bad grammar and punctuation can really fsck up the comprehension of a story?
Please learn the correct, use of the comma. This sentence does not make any sense as it is written...
"When strained in a particular manner, nanobubbles formed on a sheet of graphene, within which electrons came to occupy particular, quantum energy levels rather than the usual, continuous range of energies in unstrained graphene."
Karl's version of Parkinson's Law: Work expands to exceed the time alloted it.