The photographer does hold the copyright to the photograph, but copyright isn't a right to make copies of the photograph, it's a right to prevent people from making copies of the photograph. People other than the photographer may hold other rights that also preclude making copies of the photograph (but that doesn't mean they're allowed to make copies of the photograph either).
Your first link is very different from the other two. In the first case, the guy who owned the camera initially claimed he was not the photographer--specifically that a monkey picked up his camera and took the photo itself. (The photo became a bit of a sensation on this basis, but when it came to light that you don't get copyright just by owning a camera, the guy changed his story, and instead claimed he had got the shot ready, and just let the monkey push the button.) In this case, the question is about who (or what) the photographer was, and therefore who (if anyone) holds copyright (since monkeys can't hold copyright). In this case, it is questionable whether the guy who owned the camera holds copyright, and therefore whether he can preclude others from making copies.
The other two links don't relate to disputes about who the photographer is, and therefore don't relate to disputes about who holds copyright. In these cases, it is not in question that the photographer can preclude people from making copies. What is in question is whether other people hold other rights that also preclude the photographer from making copies.