I can't imagine why it would.
To the best of my knowledge, Google uses pretty much no Windows servers themselves(at least not for any of their public facing products, they almost certainly have some kicking around) and "a vast number of instances of custom in-house server applications" is among the least plausible environments for a Windows server deployment, so that is unlikely to change.
On the desktop side, Google has a bunch of stuff that runs on Windows; but it all communicates with Google's servers over various ordinary web protocols and stores local files with the OS provided filesystem. The benefits of EXT4 on Windows would have to be pretty damn compelling for them to start requiring a kernel driver install and a spare unformatted partition.
I suppose it is conceivable that some Google employee might decide to do it, for more or less inscrutable reasons; but it would have no connection at all to Google's broader operation or strategy.